SuperSeven747
I’m sorry but I think you are over thinking things.
The fact that an air- vent tube is fitted alongside the large bore filler tube is surely evidence that it vents to the outside, somewhere within the filler/cap system.
The above photo by @Johnny Rocket shows the inlet tube opening into the filler body ,presumably to the side of the shelf that the rubber seal on the cap seals to.
The failure to vent is probably down to a number of possible causes.
1. Oversized rubber seal on locking cap.
2. Malposition of the rubber seal.
3. Obstruction of the vent orifice.
View attachment 106814
That is an air breather pipe, and allows air to EXIT as you fill the water tank.
When the cap is closed, that pipe does not vent to the outside air (as the head of that pipe joins the plastic housing BEHIND the filler cap, so it can't be used for drawing in air when water is used and the cap closed.